Centrosaurus brinkmani Ryan & Russell, 2005 Coronosaurus is a genus of centrosaurine ceratopsian dinosaurs which lived in the Late Cretaceous, in the middle Campanian stage.
Later studies, however, did not recover a monophyletic clade with the genus' type species Centrosaurus apertus in phylogenetic analyses.
[1] The specific name brinkmani honors Donald Brinkman, for his research in palaeoecology of the Late Cretaceous environments of Alberta.
[3] It had as an adult inflated supraorbital horncores — the "brow horns" above the eye sockets — but not elongated as in Zuniceratops, chasmosaurines, and more basal centrosaurines (like Albertaceratops and Diabloceratops), that project laterally (to the sides) over the orbit.
They develop through ontogeny, the growth of the individual, as short spines that may fuse along their adjacent margins into larger, irregular bone masses.
For example, the nasal of Coronosaurus closely resembles that of Centrosaurus apertus in its unfused juvenile or subadult and fused adult forms and appears to have undergone a similar ontogenetic changes.
All juvenile and most adult specimens have gently recurved anterior and posterior margins resulting in most horns having an apex that is oriented at least slightly caudally (backwards).
[2] In its original description, Ryan & Russell (2005) considered Coronosaurus to represent a new species of Centrosaurus on the basis of a small phylogenetic analysis.
They found C. brinkmani to be the sister taxon of the clade of Styracosaurus and C. apertus, while Pachyrhinosaurus and other derived centrosaurines were in a separate lineage.
Their revised analysis had significantly better resolution than that presented by Farke et al. (2011), due in part to the additional scoring of missing characters for some taxa based on direct observation of their specimens.
[4] The cladogram presented below follows a phylogenetic analysis by Chiba et al. (2017):[9] Diabloceratops eatoni Machairoceratops cronusi Avaceratops lammersi (ANSP 15800)