Danuvius guggenmosi is an extinct species of great ape that lived 11.6 million years ago during the Middle–Late Miocene in southern Germany.

[1] Danuvius thus had a method of locomotion unlike any previously known ape called "extended limb clambering," she says, walking directly along tree branches as well as using arms for suspending itself.

Like those of other dryopithecines, the molars of Danuvius were wide, and there was a broad length between the two cusps; however, the premolars had three roots instead of two, and the canines were more vertically oriented rather than somewhat sticking out.

This may have caused lordosis (the normal curvature of the human spine) and moved the center of mass over the hips and legs, which implies some habitual bipedal activity.

[1] The robust finger and hypertrophied wrist and elbow bones indicate a strong grip and load bearing adaptations for the arms.

Danuvius was likely able to achieve a strong grip with its big toes, unlike modern African great apes, which would have allowed it to grasp onto thinner trees.

Extended limb clambering emphasizes knee extension and lordosis, as well as the suspensory mechanisms seen in apes, and may be a precursor to obligate bipedalism seen in human ancestors.

The late Miocene may have been the beginning of a drying trend characterized by increased seasonality, causing deciduous forest to turn into a less dense woodland, and fruit and leaf production to occur cyclically rather than year-round.

[9] The late Miocene cooling trend may have led to the replacement of more tropical flora by mid-latitude and alpine varieties, and ultimately the extinction of European great apes.