[1][2][3] This is allocated in the East-Iberian area, that during the Middle Jurassic was part of a Carbonate platform system, influenced by tectonic activity and fault lines, along the Iberian and Catalan Coastal mountain ranges of Spain, with an exposure of up to 500 km.

[1][7] The El Pedregal Formation lithology is dominated by mudstone and wackestone limestones with fine sediments, including microfilaments, echinoderm fragments, and pellets, with less important sequences with interbedded marls, which are indicative of a low-energy marine environment.

[12] Conversely, deeper, sediment-starved areas feature "type 1" taphonic populations, characterized by juvenile, undamaged ammonites within homogeneous molds, typical of condensed deposits from transgressive phases.

[2] Due to be located adjacent to an isolated island, the Camarena locality insect biota likely wasn't too specialized, with generalists more likely to adapt to these environments and inflict similar damage.

[2] Modern equivalents capable of leave similar patterns in extant cycadophytes include caterpillars from genera like Eumaeus (Lycaenidae) and Chilades, along other lepidopteran families, such as Tineidae, Nymphalidae, and Erebidae.