Umbrella cells form the outermost layer of the urothelium, which is a special type of epithelium found in the renal pelvis, the ureters, and the urethra.
Umbrella cells are special in that they can contain multiple nuclei.
Their apical membrane contains numerous invaginations, which allows the cells to stretch during urination.
[3] These increase the strength of the urothelium.
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