[2] Although the experiment is inaccurately known as the Blackman Reaction, a significant part of the work was carried out by Matthaei and the 1904 paper communicated to the Royal Society by Francis Darwin bears only her name.
[3] In 1905, she married Albert Howard, imperial economic botanist to the government of India.
They argued that plants should be studied in the context of their habitat and that food grown in humus-rich soil would be beneficial to health.
Beginning in 1924, they oversaw the planning and construction of the Institute of Plant Industry at Indore.
She suddenly died in Genoa shortly before their planned retirement and return to England.