Glomerida

GlomeridaeGlomeridellidae Trachysphaeridae (=Doderiidae) Plesiocerata Verhoeff, 1910 Glomerida is an order of pill-millipedes found primarily in the Northern Hemisphere.

Like the Sphaerotheriida (so-called Giant Pill-millipedes), they are capable of enrolling into a ball ("volvation"), a trait also shared with the unrelated pillbugs (Oniscidean crustaceans).

The head is relatively large and round, possessing long, slender antennae, and horseshoe-shaped Tömösváry organs.

[7] Numerous undescribed fossils are known from the Cenomanian aged Burmese amber, which constitute the oldest known records of the order, though the group probably originated much earlier.

Oniscomorphs are united with the poorly known Glomeridesmida in the infraclass Pentazonia, which is characterized by posterior telopods and relatively short bodies, and is the sister group to the Helminthomorpha, or "worm-like" millipedes.

[14][15] However, some studies based on DNA sequence comparisons have suggested that Glomerida is more closely related to Glomeridesmida than to Sphaerotheriida,[15][16] a hypothesis which would imply the enrolling behavior evolved twice or was lost in the ancestors of Glomeridesmidans.

Male Glomeris punica from Tunisia. The enlarged rear legs are the telopods
Trachysphaera pyrenaica