Gnathichnus

Gnathichnus is a trace fossil on a hard substrate (typically a shell, rock or hardground made of calcium carbonate) formed by regular echinoids as they scraped the surface with their five-toothed Aristotle's Lantern feeding structures.

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Oral surface of Strongylocentrotus purpuratus showing the teeth of the Aristotle's Lantern, which can make the trace Gnathichnus .