Not much is known about the Negative Syndicates (ネガティブシンジケート, Negatibu Shinjikēto) other than that they are individual organizations whose respective leaders, foot soldiers, and monsters try to steal the Precious and use them for evil deeds.

The Jyaryu Clan (ジャリュウ一族, Jaryū Ichizoku) are the second of the Negative Syndicates who seek to destroy mankind and make the Earth more suited to their reptilian bodies.

The hand-held Accellular henshin devices use their half-open Scan Mode to determine the "Hazard Level" of each relic; the higher the number, the more important or dangerous the Precious is.

They looked back at each of the 29 previous teams, referencing various items within the older shows, and later on, wearing the costumes and using other props from the series, such as the "civilian" clothing.

A mysterious being called Muse (ミューズ, Myūzu) awakens, announcing the Precious to whoever gets to her first, causing a 5-way confrontation between the Boukengers, Gajya, the Jyaryu, Dark Shadow and the Questers.

The villain for this special, Time Demon Chronos (時の魔神クロノス, Toki no Majin Kuronosu), abducts all of the Boukenger, save for Eiji, into an alternate dimension.

They then fight Chronos who has purchased Goodomu Engines from Gajya to bring back Furabiijo from Hurricaneger, revive Duchess Org TsueTsue from Hyakujuu Sentai Gaoranger and Sorcery Priest Meemy of Magiranger.

While the remaining Boukengers are in the other dimension, they meet up with Hikaru (Magi Shine of Magiranger) and Smoky, his cat genie, and try to find a way to escape.