It is a descendant of macrohaplogroup N. Haplogroup S mtDNA evolved within Australia between 64,000 and 40,000 years ago (51 kya).
Haplogroup S2 found in Willandra Lakes human remain WLH4 dated back Late Holocene (3,000–500 years ago).
[3][4] The following table lists relevant GenBank samples: This phylogenetic tree of haplogroup S subclades is based on the paper by Mannis van Oven and Manfred Kayser Updated comprehensive phylogenetic tree of global human mitochondrial DNA variation[1] and subsequent published research.
The TMRCA for haplogroup S is between 49 and 51 KYA according to Nano Nagle's Aboriginal Australian mitochondrial genome variation – an increased understanding of population antiquity and diversity publication that published in 2017.
[2] Phylogenetic tree of human mitochondrial DNA (mtDNA) haplogroups This genetics article is a stub.