[2] At the beginning of the film, Leo is an adult and learns that his mate, Lyra, has just given birth to twin cubs: Lune (pronounced Lu-Ney) and Lukio.

Minus seek the help of Ham Egg to lead them to the source of the Moonlight Stone so it can be salvaged.

Lemonade are shocked by Ham Egg and his "friends", who lack concern for any form of animal.

Eventually, a stork from the jungle finds Lune, tells him of how the disease is killing off many animals, and she leaves with her flock.

Although Lukio comes down with the same ailment, Dr. Moustache is able to cure her and others, including Pagoora's son thus proving humans can be friends to animals.

They go up the mountain, guided by Earth Mother who helps them fight a pack of dire wolves.

The film ends with Lune returning from the human world, and Dr. Moustache taking the cub to see his father's pelt.

He was the most courageous soul I've ever met, a true king who gave his life to save the land."