Marmyan

Burmese amber has been radiometrically dated using U-Pb isotopes, yielding an age of approximately 99 million years, close to the Aptian – Cenomanian boundary.

[1] Marmyan is one of three Burmese amber coccid genera that Koteja described in the same paper, the other two being Burmacoccus and Albicoccus placed into the monotypic families Burmacoccidae and Albicoccidae respectively.

The extinct genera Albicoccus, Apticoccus, Grimaldiella, Kukaspis, Marmyan, Palaeosteingelia, Palaeotupo, Solicoccus, Turonicoccus and possibly Pennygullania, all with simple rows of eyes, were found to form a clade that included Burmacoccus.

In contrast to the work of Hodgson and Hardy, the 2015 analysis recovered Marmyan as a neoccoid genus sister to the family Eriococcidae, as was suggested by Koteja.

The peniel sheath is divided into a basal rounded section and a narrow style, slightly downward curved, with pointed end.