It covers an area of 307,720 km2 in all of Hungary, most of Slovakia, about half of Croatia and Slovenia, around a third of Bosnia and Herzegovina, Romania, and Serbia, and minor parts of Austria, Czech Republic, and Ukraine.

The plant communities include mixed oak-hornbeam forests, azoal floodplain vegetation and lowland to montane herb-grass steppes.

Sub-Mediterranean thermophilous bitter oak forests grow in warmer areas.

Higher elevation areas are dominated by European beech and silver birch, downy birch, European aspen and sometimes by conifers Abies alba, Picea abies, Pinus sylvestris, Pinus nigra.

Lowland to montane herb-grass steppes dominate large areas of the ecoregion.