Colonies are snowy white, circular, and convex with smooth surfaces and entire edges.
[4] Tests for catalase, oxidase, urease, deaminase, and lipase are positive.
Amylase, protease (gelatin), tryptophanase (indole), nitrate reductase, cysteine desulfurase, and agarase tests are negative.
[4] The following carbon sources are utilized: acetate, lactate, malate, fumarate, pyruvate, propionate, citrate, succinate, oxaloacetate, butyrate, 2-oxoglutarate7 glucose, glycerol, sorbitol, DL-alanine, DL-glutamate, DL-proline, DL-aspartate, and DL-asparagine.
[4] The following carbon sources are not utilized: maltose, D- fructose, sucrose, lactose, D-xylose, D-ribose, formate, cellobiose, D-mannose, L-fucose, melibiose, melezitose, L-rhamnose, sorbose, trehalose, methanol, ethanol, erythritol, propanol, benzoate, malonate, DL-arginine, glycine, DL-serine, DL-isoleucine, DL-lysine, DL-histidine, DL-ornithine, DL-methionhe, DL- valine, DL-threonine, and DL-tryptophan.