In 2012, following a comprehensive review of the dileptid group, it was moved to the genus Pseudomonilicaryon, which is distinguished from Dileptus by the shape of its macronucleus.

The macronucleus of Pseudomonilicaryon is arranged like a string of beads, whereas that of Dileptus is composed of small nodules scattered throughout the cytoplasm of the cell.

A rapacious predator, described by one writer as "surely the king of beasts of the Protozoa," it feeds exclusively on other microscopic organisms.

When the proboscis makes contact with its prey, toxic extrusomes are released, disabling the smaller organism, which is then gathered into the dileptid's cytostome.

Smaller prey are swept into the mouth by the movement of cilia, while large organisms are pushed toward the oral bulge by "the writhings of the proboscis.