In the era of commercial typesetting in metal type, they were usually manufactured in strips of 6-point lead.
Objects (including text frames) positioned in the slug area are printed but will disappear when the document is trimmed to its final page size.
[1] Slugs, or slug lines, are also the name for incidental typeset lines of type that are intended either for the printer's or binder's benefit (such as a collation mark, a catch line, or a galley slug) or as advertising for the producer of the printed piece (such as a line of type showing the name of the printer, the printer's item number or job number, and the telephone number of the printer in order to make reorders simple).
This term is also used in web publishing to refer to short article labels that may be used as part of a URL.
Slugs are usually derived from an article's title and are limited in length, and to a specific set of characters (to prevent percent-encoding); often only letters, numbers, and hyphens are allowed.