The hour-long episode brings to a climax the romantic character arc between Niles and Daphne, a significant running plotline for the first seven years of the show's production.
Frasier, Daphne, Niles, Mel, and Martin return from the funeral of Morrie, the doorman of Elliott Bay Towers.
Martin bears a final gift from Morrie; a rare bottle of wine retrieved from France following the Second World War.
Mel leaves to prepare for a mid-week retreat to celebrate her first six months with Niles, while he confesses to Frasier that he feels anxious about their relationship.
After Niles denies that he still possesses lingering feelings for Daphne, Frasier urges his brother to seize the moment and move on from his habitual sense of caution.
Frasier confesses that he does not believe either Daphne or Niles when they claim to be happy, leading Martin to warn him about interfering; two marriages are at stake.
The next morning, as the wedding takes place, Niles sits in the front cabin of the Winnebago, unable to watch Daphne be married.