Tapejaridae

Torukjara bandeirae, Caiuajara dobruskii, Tupandactylus imperator, Tapejara wellnhoferi, Huaxiadraco corollatus, and Sinopterus dongi Tapejaridae (from a Tupi word meaning "the lord of the ways") is a family of azhdarchoid pterosaurs from the Cretaceous period.

[6] Tapejarids had unusually reduced shoulder girdles that would have been slung low on the torso, resulting in wings that protruded from near the belly rather than near the back, a "bottom decker" arrangement reminiscent of some planes.

[8][9] Tapejaridae was named and defined by Brazilian paleontologist Alexander Kellner in 1989 as the clade containing both Tapejara and Tupuxuara, plus all descendants of their most recent common ancestor.

[16] Later, in 2006, British paleontologists David Martill and Darren Naish followed Unwin's concept, and provided a revised definition for Tapejaridae was also proposed: the clade containing all species more closely related to Tapejara than to Quetzalcoatlus.

[20] In 2020, in the description of the genus Wightia, an opposing subfamily was named, Sinopterinae, consisting of tapejarids more closely related to Sinopterus than Tapejara.

Therefore, they favored the term Thalassodromidae to have consistency with other studies that used the same name, despite finding them to form a natural grouping with Tapejaridae in their phylogenetic analysis (per the Kellner model).

[22] In 2024, Pêgas rejected this redefinition of Tapejaridae in light of non-compliance with phylocode rules, applying the Tapejara and Sinopterus definition and deeming Eutapejaria a synonym.

[24] Thalassodromeus sethi Kariridraco dianae Tupuxuara leonardii Tupuxuara longicristatus Caupedactylus ybaka Aymberedactylus cearensis Eopteranodon lii Huaxiapterus benxiensis Huaxiapterus corollatus Sinopterus dongi Tapejara wellnhoferi Europejara olcadesorum Caiuajara dobruskii Tupandactylus imperator Below are two cladograms representing different concepts of Tapejaridae.

Additionally, his analysis further differs from that of Andres in finding both Tapejaridae and Thalassodromidae within Tapejaromorpha, which corroborates the close relationship between thalassodromids and tapejarids, similar to Kellner.

Reconstructed profiles of three Brazilian species; from top to bottom, Tapejara wellnhoferi (A), Tupandactylus navigans (B), and Tupandactylus imperator (C)
Skeletal reconstructions of Thalassodromeus , phylogenetically controversial pterosaurs classically assigned to Tapejaridae
Reconstructed skull of Caupedactylus , a pterosaur recovered either within Tapejarinae or just outside Tapejaridae in its own clade, Caupedactylia