[2] It had a long neck and tapir-like, three-toed feet,[2][3] and like other litopterns and modern horses, tapirs and rhinos, it bore its weight on its middle toes.

There are either ten or seven species, and in 2014 Schmidt and Ferrero put forward that the genus needed a full taxonomic revision due to this issue.

[4] In 1931, fossils found in the late Miocene Arroyo Chasicó Formation were assigned to the genus under the name Theosodon hystatus.

[4] Proadiantus Tricoelodus Polymorphis Pternoconius Coniopternium Cramauchenia Theosodon Scalabrinitherium Oxyodontherium Huayqueriana Paranauchenia denticulata Paranauchenia hystatus Cullinia Promacrauchenia Windhausenia Xenorhinotherium Macraucheniopsis Macrauchenia Theosodon was a terrestrial and cursorial animal that may have lived in both forests and more open environments.

Some modern animals such as giraffes feed on thorny plants, and their nostrils are also further back and face more upwards than forwards.