The younger Toudouze wrote on such topics as art, architecture, travel and French naval history.
Although he penned numerous adventure novels and short stories, he is today remembered for a single work: "Three Skeleton Key".
The men escape into the lighthouse gallery, which has a metal trapdoor that the rats cannot gnaw through, and are able to use the light to signal an investigating patrol boat.
Eventually the rats are lured off the island onto a barge loaded with meat, which is then set on fire by incendiary shells.
The fates of the men are then revealed by the narrator: Le Gleo went insane from the events and was locked away in a French asylum, and Itchoua dies of infection from rat bites and scratches.