Wynne was the youngest son of Robert Wynne (a canon of Bangor Cathedral and rector of Llanddeiniolen and Llaniestyn) and his wife Catherine Madryn, heiress of Llannerch Fawr.
William Wynne was born before 12 November 1671 and educated at Jesus College, Oxford (like his father and elder brother).
Whilst at Oxford, he was part of the circle of Edward Lhuyd, the Welsh naturalist, botanist, linguist, geographer and antiquary.
Wynne's version, which was reprinted in 1702 (unchanged), 1774 and 1812 (with changes each time), and in 1832 (with topographical notes), was the starting point in English on medieval Welsh history for nearly two hundred years.
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