Zeus (aliases Dmel\Rcd-1r, Rcd-1 related/Rcd-1r, CG9573, Drcd-1r/Drcd-1 related)[1] is a male fertility gene in Drosophila,[2][3][1] known especially from D. melanogaster[1] and D.
[2] Zeus arose via a retrotransposition from Caf40,[3] a DNA-binding protein.
[2] It differs by 107 amino acid substitutions.
[2] It retains approximately 30% of Caf40's DNA-binding sites but over the past perhaps 4-6 million years has acquired another 193.
[2] These new binding actions regulate hundreds of downstream reproduction related genes.