After training in the United States with Consolidated B-24 Liberators, it deployed to the European Theater of Operations, where it engaged in the strategic bombing campaign against Germany.

The aircrews ferried their planes under the control of Air Transport Command via the southern route from Florida through Puerto Rico, Brazil, Senegal, and Morocco to England.

The 707th was part of the first United States Army Air Forces group to complete the Transatlantic hop from Brazil to Africa without the installation of additional bomb bay fuel tanks.

It supported Operation Overlord, the invasion of Normandy in June 1944 by attacking strong points, bridges, airfields, transportation, and other targets in France.

The squadron aided ground forces at Caen and Saint-Lô during July by hitting bridges, gun batteries, and enemy troops.

It flew low level missions to drop medical supplies, arms, and food to airborne and ground troops near Wesel during Operation Varsity in March 1945.

[4][d] President Truman’s reduced 1949 defense budget required reductions in the number of units in the Air Force,[13] as a result, the 707th was inactivated in March 1950.