Alan Garen was an American geneticist who co-discovered suppressor mutations for tRNA.
The Garen lab also showed that certain triplet codons (5'-UAG, 5'-UAA, and 5'-UGA) failed to bind amino acids.
[1] Thus, the Garen lab and Brenner labs are both credited with discovery of the stop codons of the genetic code.
[2] Garen was a professor at Yale University between 1963 and 2021.
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