Ancient Southern East Asian

[3] In the mid-Holocene, southward migrations of millet farmers from the Yellow River harboring ANEA ancestry (and also to lesser degree a reverse geneflow of ASEA rice farmers from the Yangtse River to the north) resulted in the coastal East Asian ancestry cline that exists to this day.

This technological spread was a result of the migration of southern East Asian agriculturalists that carried ASEA ancestry.

[6][5][7] Ancient DNA of first farmer individuals from Mainland Southeast Asia dated at c. 4kya derives most of its ancestry from the ASEA lineage, with significant admixture from a local hunter-gatherer population.

Hence, the first spread of farming in Mainland Southeast Asia is widely assumed to be linked to the expansion of the Austroasiatic languages.

The rapid maritime expansion of the early Austronesians starting c. 5,000–4,000 years ago brought ASEA ancestry from Taiwan to the Philippines, the Indonesian archipelago and Oceania, initially with little admixture from local populations, as can be seen from 2,900–2,500 year-old Lapita-related individuals from Vanuatu and Tonga,[14] and from ancient 2,800–2,200 year-old DNA of the first settlers of Guam.