In his early years, which I covered in the X-Factor Forever miniseries... Apocalypse encountered the Celestials and realized there was a time when humanity might be judged unworthy and destroyed.

"[6] Guice admitted to difficulty recalling the details behind redrawing the last page of issue #5: "The best I can remember now is putting his look together pretty much right on the pencil page—just adding bits of costuming business which hinted toward his true appearance when we'd eventually see him in full reveal.

During his run on Cable, Robert Weinberg planned a story to reveal that Apocalypse was the third Summers brother, a mysterious sibling to the mutants Cyclops and Havok.

Knowing who En Sabah Nur is fated to become and where he is, Rama-Tut sends his General Ozymandias and an army to destroy the Sandstormers and find the young Apocalypse.

Vowing revenge on Rama-Tut, En Sabah Nur enters the Pharaoh's city posing as a slave and draws the romantic attention of Ozymandias's sister, Nephri.

Rama-Tut flees and En Sabah Nur uses the Celestial technology to transform his former tormentor Ozymandias into a blind clairvoyant made of living stone, now enslaved to Apocalypse.

[18] As the millennia pass, Apocalypse travels around planet Earth, convincing civilizations that he is a deity (inspiring different myths as a result) and manipulating them into fighting wars.

Apocalypse encounters the near-immortal human offshoot race known as Eternals, primarily the members Ikaris and Sersi, who refer to him as their "Ancient Nemesis".

In AD 1013, Apocalypse seeks to destroy the Asgardian Thor, who he knows will cause him trouble in the future, according to information obtained from Rama-Tut,[21] yet the plans are foiled by Odin.

In the 12th century, Apocalypse encounters the Eternal Sersi again while awakening latent mutant powers in a crusader named Bennet du Paris, also known as Exodus.

Although Apocalypse is severely defeated, the young Nathan is infected with a techno-organic virus and is sent to the future with a woman named Askani to be cured.

After a long healing slumber, Apocalypse, fully restored, awakens with Ozymandias at his side and quickly learns of the present danger: Onslaught.

[41] He observes the conflict between the psionic entity and Earth's heroes with Uatu the Watcher, who suggests to Apocalypse a course of action: an alliance with the one who hated him the most, Cable.

To test this newest recruit, Apocalypse set War against the New World Order, a shadow cabinet organization that intends to conquer the planet.

Reappearing inside a Sphinx-shaped ship, Apocalypse confronts the X-Men with his newly assembled cadre of Horsemen on the front lawn of the X-Mansion.

Upon learning of Apocalypse's return, X-Force seeks to kill him, but when they discover he is a child, Psylocke decides to protect him, believing they can rehabilitate him and train him as a force for good.

The boy, code-named Genesis, helps X-force fight Archangel and when the battle is over, Fantomex enrolls him in the Jean Grey School for Higher Learning.

After Deadpool's failed attempt to rescue Evan, the boy dons the Celestial armor to prevent Wolverine's death at Daken's hands and nearly kills Brotherhood members Sabretooth and Mystique.

[72] In the energy released when the Scarlet Witch and Doctor Doom use the combined power of order and chaos spells to suppress the Red Skull's personality in favor of the sliver of Professor X's that still exists in his brain, Evan ascends to adulthood, stepping out of the rubble with a looming, muscular body and the presence and appearance of Apocalypse.

When Havok and Cyclops argue with Steve Rogers and the new Captain America over who should take custody of the body of Red Skull, Evan convinces the Summers brothers that Professor X is dead and they have other problems to deal with.

His efforts are not in vain, as he begins to impose his own consciousness onto a human test subject, the fourth attempt, lending it his own regenerative powers to withstand the assault.

Once the wave subsides, Apocalypse finds himself in a strange new land that he deduces was the result of the explosion within the Celestial machine that must have caused a dimensional rift which threw him into another dimension.

Apocalypse then gave his perfect host body a trial run as he killed the remaining test subjects, while reducing the entire island to rubble that was reclaimed by the sea.

[86] During "Dawn of X", Apocalypse accepts Xavier's invitation to enter Krakoa[87] and even is among the Quiet Council, a group of fourteen powerful and experienced mutants who serve as the island's lawgivers.

[89] Its also revealed that at an uncertain point in the distant past apparently before En Sabah Nur merged himself with the Celestial technology, he met and married his wife Genesis.

As time passed he and Genesis conceived four children, two sons and two daughters who eventually became their First Horsemen when the island was split by a force wielding the Twilight Sword.

In the end, Genesis, the Horsemen, and Arakko decided to move to Amenth in order to hold off the Amenthi forces and deny Annihilation access to Earth by sealing the chasm, while leaving Apocalypse behind, since he wasn't strong enough, much to his and Krakoa's sorrow.

However, before Genesis went to Amenth, she made Apocalypse promise to find and judge the succeeding mutants in following years in order to help them fight the threat.

[106] In the 2010 "Heroic Age" storyline, versions of Apocalypse and his Horsemen from a possible future appear in the Avengers Tower after Kang breaks time itself.

Cable makes the most concrete implication that evolving is the major element of his powers when he remarks that Xavier must kill Apocalypse quickly before he adapts to his attacks and becomes immune to the telepathy.