D. artesiana (Danzé-Corsin) D.Edwards (2006)[1] Danziella is a genus of extinct vascular plants of the Early Devonian (around 420 to 390 million years ago).
Fossils found in the Artois region of northern France were first described as Zosterophyllum artesianum,[2] but a later review by Edwards showed that they did not fit the circumscription of that genus.
Spore-forming organs or sporangia were borne on all sides, spaced irregularly on stalks up to 3 mm long which held them horizontally, and not forming a distinct 'spike'.
[3] In 1956 Danzé-Corsin constructed a new species, Z. artesianum to accommodate this plant, arguing that it resembled Z. llanoveranum but had its sporangia arranged in a much laxer spike.
[4] Edwards agrees that Danziella may be part of the stem group of the zosterophylls and hence the lycophyte clade, but says that it could also belong elsewhere and show convergent evolution of the mechanism by which the sporangia split to release spores.