The flocculus (Latin: tuft of wool, diminutive) is a small lobe of the cerebellum at the posterior border of the middle cerebellar peduncle anterior to the biventer lobule.

[4] The climbing fibers then send the image or projection to the part of the brain that receives electrical signals and generates movement.

Finally, pontocerebellar projections carry vestibulo-occular signals to the contralateral cerebellum via the middle cerebellar peduncle.

The flocculus is a part of the vestibulo-ocular reflex system and is used to help stabilize gaze during head rotation about any axis of space.

Neurons in both the vermis of cerebellum and flocculus transmit an eye velocity signal that correlates with smooth pursuit.

The flocculus appears to be included in a VOR pathway that aids in the adaptation to a repeated shift in the visual field.

[9] Bilateral lesions of the flocculus reduce the gain of smooth pursuit, which is the steady tracking of a moving object by the eyes.

Instead, the bilateral lesions of the flocculus result in saccadic pursuit, in which smooth tracking is replaced by simultaneous rapid movements, or jerking motions, of the eye to follow an object toward the ipsilateral visual field.

These lesions also impair the ability to hold the eyes in the eccentric position, resulting in gaze-evoked nystagmus toward the affected side of the cerebellum.

Some symptoms of common vestibular diseases include: head tilting, an inability to stand, ataxia, dizziness, vomiting and strabismus.