Foundation species

A foundation species can occupy any trophic level in a food web (i.e., they can be primary producers, herbivores or predators).

[3] A study conducted at the McKenzie Flats of the Sevilleta National Wildlife Refuge in New Mexico, a semiarid biome transition zone, observed the result of loss of a variety of different dominant and codominant foundation species of plants on the growth of other species.

[4] Another study observed the effects of loss of foundation eastern hemlocks (Tsuga canadensis) in a forest ecosystem.

This study observed the effects that a loss in eastern hemlocks would have on the populations of arthropods, such as ants, beetles, and spiders, since these species are known indicators of environmental change.

It was found that species at the base of the food web are less strongly, and carnivores are more strongly facilitated in foundation species' food webs than predicted based on random facilitation, resulting in a higher mean trophic level and a longer average chain length.

[11][12][13][14][15][16] This form of non-trophic facilitation by foundation species has been found to occur across a wide range of ecosystems and environmental conditions.

[17][18] In harsh coastal zones, corals, kelps, mussels, oysters, seagrasses, mangroves, and salt marsh plants facilitate organisms by attenuating currents and waves, providing aboveground structure for shelter and attachment, concentrating nutrients, and/or reducing desiccation stress during low tide exposure.

Californian forest of giant kelp , a foundation species [ 1 ]
Tsuga canadensis
Foundation species enhance food web complexity
In a 2018 study by Borst et al ...
(A) Seven ecosystems with foundation species were sampled: coastal ( seagrass , blue mussel , cordgrass ), freshwater ( watermilfoil , water-starwort ) and terrestrial ( Spanish moss , marram grass ).
(B) Food webs were constructed for both bare and foundation species-dominated replicate areas.
(C) From each foundation species structured-food web, nodes (species) were randomly removed until the species number matched the species number of the bare food webs.

The presence of foundation species strongly enhances food web complexity, facilitating particularly species higher in the food chains. [ 7 ]