This remarkable potential makes them useful for basic research on the function of the human body, for drug discovery and testing, and as a source of cells and tissues for transplantation medicine.

[4][2] Science later featured induced pluripotent stem cells in its “Scientific Breakthrough of the Year” article, 2008.

His doctoral thesis involved understanding genetic imprinting in early mammalian development under the mentorship of Davor Solter at the Wistar Institute.

He joined the Wisconsin Regional (now National) Primate Research Center on campus as its chief pathologist in 1995.

[7] He is also a founder of Cellular Dynamics International, a Madison-based company producing derivatives of human induced pluripotent stem cells for drug discovery and toxicity testing.