[1] He attended San Jose State University for his undergraduate, finishing with a BA in psychology and physics in 1979.
[2] He went on to receive his PhD at the University of California, Santa Barbara in 1986 under the direction of Gerald Jacobs.
[3] After his PhD, he stayed at the same institution as a postdoctoral researcher for several years before starting a permanent position at the Medical College of Wisconsin.
Their work on treating color-blindness in monkeys received some attention in the popular science press.
In this work, they gave gene therapy to two red-green color-blind squirrel monkeys, combined with training.