Marginal distribution (biology)

In some cases, geographical range limits are entirely predictable, such as the physical barrier of an ocean for a terrestrial species.

This is based on the assumption that the habitat is most ideal at the centre of a distribution and ecological conditions decline towards the margin.

Reduced gene flow between central and peripheral populations also limits the genetic diversity at the margins.

High selection pressure, due to a less than ideal habitat at the margin, furthermore reduces genetic diversity.

In Europe, for example, the geographical spokes sticking out of the continent in the south - the Iberian Peninsula, Italy and the Balkans served as refugia for warmth-adapted species during the Ice Ages.

This is often seen in organisms with high water demands, whose survival and reproduction is limited by dry conditions.

[15] There are many other abiotic factors that can determine a species range, including dissolved oxygen, conductivity, alkalinity and pH.

Where two similar species share an overlapping range, competition often causes the distributions to shift to exclude one of the two.

[24] Humans have also initiated many range expansions by introducing species to new locations both intentionally and accidentally.

[25] In most cases combinations of factors are responsible for limiting the geographic range edge of species.

The pine moth, Thaumetopoea pityocampa , expanded its range northward in France.
Individuals of the freshwater gastropod Lymnaea stagnalis were found to have higher occurrences of trematode parasites at the edge of their distribution as compared with those of the core.
The bronzed cowbird , Molothrus aeneus , continues to expand its range in New Mexico and Texas due to deforestation .