It is the tract of the mossy fiber projections, both inhibitory and excitatory from the granule cells of the dentate gyrus.
The interneurons of the stratum lucidum are generally found to be local circuit neurons remaining within the CA3 region.
This situation is described in the mossy fiber axon connection in the CA3 stratum lucidum region of the hippocampus as is in relation to Purkinje cells.
Spiny neurons are of importance because of their "pivotal role in motor control, habit formation, and motivated behavior".
The somata of aspiny neurons are for the most part bipolar, generating 2–5 primary dendrites "that to a varying extent displayed varicose swellings in their course".
[3] This article incorporates text in the public domain from the 20th edition of Gray's Anatomy (1918)