Testudo hellenica

Testudo hellenica is an extinct genus of tortoise of the genus Testudo from the Miocene (Vallesian) Nea Messimvria Formation (Zone MN 10) of Greece.

[1] T. hellenica is the earliest known crown-Testudo from Greece (according to Garcia et al., 2020), since the next oldest Testudo species, T. marmorum, from Greece come from the Turolian (7.3-7.2 ma) Pikermi beds.

[5][6][7] In order to observe the detail of the carapace, the specimen was completely prepared.

[1] The holotype, LGPUT RPI-216, preserves a carapace, plastron and several bones underneath, including a fused humerus and scapula.

[1] Since the domed pygal plate curves inwards, this indicates that the holotype individual was a male.