The story centers on an American man living in Britain, Ian Stone (Vogel), who is killed each day by mysterious beings.
Late one night while driving home from a painful loss on the ice, Ian thinks he sees a dead body near the railroad tracks.
Investigating the grisly discovery, Ian is attacked by the "corpse", forced onto the tracks and run over by a train.
As they seek refuge on a subway train heading out of town, Jenny confesses to her memories of Ian's former lives.
When they disembark they are again confronted by a group of Harvesters, each a mass of pulsating veins, pitch-black muscles and flesh.
Some of the barely human faces are recognizable from Ian's previous lives, including Medea who reveals a shocking truth: Ian was one of these monsters until he rebelled against the colony and managed to kill one other Harvester, and Medea used to be his "mate".
Making this connection to a human being changes the specific nature of a Harvester, which allowed the old man (and now Ian) to kill the members of their kind.
Ian regains his true power: his Harvester-self is finally born out of his human flesh, uniquely exuding goodness rather than malevolence.
Jenny has no recollection of their harrowing adventure; but Ian, having realized his abilities, will begin to take the fight to the Harvesters.