Titanoceratops

It was a giant chasmosaurine ceratopsian that lived in the Late Cretaceous period (Campanian stage, about 75 million years ago[1]) in what is now New Mexico.

Longrich believed that unique features found in the skull reveal it to have been a close relative of Triceratops, classified within the subgroup Triceratopsini.

[2] In total, the amount of material assigned to Titanoceratops means it is quite well known, along with genera like Triceratops, Vagaceratops, Pentaceratops, Chasmosaurus, Centrosaurus, Styracosaurus, and Anchiceratops.

The holotype specimen consists of most of the fore and hindlimbs, some vertebrae, a fairly complete skull with only one small section of the frill, and partial lower jaws.

[3] The specimen was later reinterpreted as a member of the Triceratopsini, the group including Triceratops, by Nicholas R. Longrich and given the name Titanoceratops ouranos in 2011.

The name Titanoceratops is derived from the Greek Titan, a mythical race of giants, keras (κέρας), meaning "horn", and ops (ὤψ), "face".

Until the Longrich's re-interpretation of Titanoceratops, Eotriceratops was thought to be the oldest known triceratopsin, and only dated to 68 million years old, from the uppermost region of the Horseshoe Canyon Formation.

Longrich interpreted the specimen as sharing more characteristics with Triceratops and Torosaurus than with Pentaceratops, and he named a new group, Triceratopsini, to contain all of them.

They agreed with Lehman's original assessment, that the features in the specimen that appeared unique were likely due simply to advanced age and unusually large size.

The Fruitland Formation is about 100 metres (330 ft) thick, and consists of sandstones, mudstones, and abundant coals deposited in a coastal floodplain.

The Kirtland Formation, which conformably overlays the Fruitland, is approximately 600 metres (2,000 ft) thick, and made up of sandstone, siltstone, mudstone, and shale.

[2] Non-dinosaurian fauna include the fishes Myledaphus bypartitus, and Melvius chauliodous; the turtles Denazinemys ornata, Denazinemys nodosa, Boremys grandis, Neurankylus baeuri, Adocus bossi, Adocus kirtlandicus, Basilemys nobilis, Asperideretes ovatus, "Plastomenus" robustus, and Bothremydidae n. gen., barberl; the crocodylians Denazinosuchus kirtlandicus, Brachychampsa montana, Deinosuchus rugosus, and Leidyosuchus sp.

Life restoration
Skeletal reconstruction, holotype material in white, with human for scale
Beds of the upper Fruitland Formation and lower Kirtland Formation in the vicinity of the Titanoceratops holotype quarry
J. Willis Stovall holds up the humerus, 1941
Holotype restored based on Pentaceratops
Front view of the mount
The fossilized leaf of Sabalites , a Cretaceous palm, documents the wet, warm climate that prevailed in Late Cretaceous New Mexico
A fossil tree stump from the Coal Creek area where Titanoceratops was found, showing the extensive coal beds of the area