The Tyers River rises below Talbot Peak, part of the Great Dividing Range, within the Baw Baw National Park at an elevation of 1,240 metres (4,070 ft) and descends steeply.

At Tyers Junction the river is joined by the confluence of the Tyers River West Branch that drains the eastern slopes of Mount Mueller from an elevation of 1,130 metres (3,710 ft) and the Tyers River East Branch that drains the southern slopes of Talbot Peak from an elevation of 551 metres (1,808 ft).

The river flows in a highly meandering course generally south, then south by east, through the Moondarra State Park, joined by one minor tributary, before reaching its confluence with the Latrobe River west of Tyers in the Latrobe City local government area.

The 30,400 thousand cubic metres (1,070×10^6 cu ft) reservoir is predominantly used to augment domestic water supplies.

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