He started graduate training at Harvard Medical School advised by E. C. C. Lin and earned a PhD in biochemistry in 1966.
[5] During his tenure, he expanded the editorial board from 26 to more than 140 and created a second track to allow scientists who were not members of the National Academy of Sciences to submit manuscripts directly.
Francis Crick wrote in his book What Mad Pursuit: At about this time, Bill Pohl, a pure mathematician, got into the act.
He pointed out, quite correctly, that unless something very special happened, the most likely result of replicating a piece of circular DNA would be two interlocked daughter circles rather than two separate ones.
I told him in a letter that if nature did occasionally produce two interlocked circles, a special mechanism would have been evolved to unlink them.