Sphyraena bolcensis is an extinct species of prehistoric barracuda known from the Eocene.
[4] More recent studies suggest that it appears to predate the evolutionary radiation of crown group Sphyraena, and thus likely represents a stem-group barracuda.
Despite being classified in the extant genus Sphyraena, its phylogenetic affinities have never been properly studied, and it thus may not necessarily be a true member of the genus, especially given it predating the modern barracuda radiation.
[4] A fossil specimen of the extinct shark Galeorhinus cuvieri has the partially-preserved remains of an S. bolcensis in its stomach.
This is reminiscent of the preference of its modern relative, the school shark (G. galeus), for also preying on barracudas, suggesting that this predator-prey relationship has lasted nearly 50 million years.