[2] In 2012, DNA analysis and statistical techniques were used to infer that a now-extinct human population in northern Eurasia had interbred with both the ancestors of Europeans and a Siberian group that later migrated to the Americas.
[3] In 2013, another study found the remains of a member of this ghost group, fulfilling the earlier prediction that they had existed.
[4][5] According to a study published in 2020, there are indications that 2% to 19% (or about ≃6.6 and ≃7.0%) of the DNA of four West African populations may have come from an unknown archaic hominin which split from the ancestor of Sapiens (Modern Humans) and Neanderthals between 360 kya to 1.02 mya.
[10] In 2015, a study of the lineage and early migration of the domestic pig found that the best model that fitted the data included gene flow from a ghost population during the Pleistocene that is now extinct.
[11] A 2018 study suggests that the common ancestor of the wolf and the coyote may have interbred with an unknown canid related to the dhole.