Glossary of botanical terms

This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general.

Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary of leaf morphology.

For other related terms, see Glossary of phytopathology, Glossary of lichen terms, and List of Latin and Greek words commonly used in systematic names.

cormose, cormouspl.

cortexes or corticesadj.

cymose Also abbreviated dicot.Also spelled disk.sing.

domatium Also aglandularAlso elliptic.

generaAlso globular.Also gramineous pl.

mycotrophic adj.

nectariferous Also spelled ochrea.Also imparipinnatepl.

opera utique oppressa pl.

phyllodineousAlso phytomelanin; adj.

rachises or rachides adj.

staminateAlso male flower.Also stipel; pl.

Often variety in common usage and abbreviated as var.Also nerve.Diminutive: virgulatepl.

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In the bud, Tetradenia riparia leaves have their upper surfaces turned toward the stem and the axil . The lower surface is abaxial ("away from the axis"), and the upper surface is adaxial .
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Welwitschia mirabilis presents an example of an acaulescent growth habit unusual in so large a plant species.
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Schematic diagrams of the accumbent arrangement of the cotyledon s and radicle in a seed of Erysimum (formerly Cheiranthus )
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Achene s on the surface of the stem of the infructescence of a strawberry
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Geranium incanum flowers are actinomorphic , having five axes of symmetry, as opposed to the two axes of symmetry of the zygomorphic flowers of most species of the related genus Pelargonium .
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Fern frond with acuminate leaflets
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Watsonia flower slit open and with one stamen bent upward to show its adnate attachment to the petal
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Diagram of a coconut fruit. The albumen (endosperm) is labelled Alb .
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Caffeine is an alkaloid with four nitrogen atoms in its carbon skeleton.
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Rothmannia leaf with extensively anastomose venation
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Androgynous flower of Sandersonia aurantiaca cut open longitudinally to show the androecium , which comprises the anther s surrounding the green central pistil
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A Neea species, family Nyctaginaceae , presents an example of an anthocarp : the calyx and style remain around the ripening fruit.
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Aphananthous flowers of oaks such as Quercus robur , being anemophilous , have no need of being conspicuous to pollinating animals.
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Apical bud of a Populus (poplar) shoot
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The apparently separate nuts of Ochrosia borbonica actually are apocarpous carpel s, two from each flower.
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Apophyses on the tips of the cone scales of Araucaria cunninghamii amount to spikes.
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Hairs on the leaves of Meniocus linifolius (formerly Alyssum linifolium ) are stellate and appressed to the leaf surface.
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Arachnoid leaves of an unidentified Gazania species
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Spine s of cactus Gymnocalycium bayrianum emerging from the areole s of the stem node s
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Seeds of a species of Blighia (ackee), one whole and one in longitudinal section, showing the pale aril
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Infructescence of wild rye, showing prominent awn s
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Anatomy of an awn and bristles on a species of the Australian grass Rytidosperma longifolium
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Axillary buds in leaf
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Barb s occur on the spines of some species of cactus, as shown here, enlarged.
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Berries of Olinia ventosa , including a cross-section showing hard seeds in the pulp
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The bifoliate compound leaves of the mopane tree, Colophospermum mopane , suggest the common name "butterfly tree".
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Cross-section of a silique of Arabidopsis thaliana , showing it to be biloculate , formed of two carpel s, morphologically a silique and not a pod
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Structure of a biternate compound leaf
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This African baobab, Adansonia digitata , has an enormous bole beneath a relatively modest canopy that is typical of this species.
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The large, colorful bract s of Bougainvillea are commonly mistaken for its petals.
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Burr s , fruits of Arctium species
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Callus tissue of Nicotiana tabacum growing on a nutrient medium in plant tissue culture
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Structure of flower of an orchid in genus Praecoxanthus , with the callus labelled
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Bearded callus of a floret of the grass species Chrysopogon filipes
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Scanning electron micrograph of the carpopodium at the base of the achene-like fruit of Zyzyura mayana , Asteraceae
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Dormant leaf buds of deciduous trees are commonly protected by imbricate cataphyll s that are shed when the bud sprouts.
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The caudex of Dioscorea elephantipes grows largely above the soil surface. Many species that form caudices grow them underground.
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Flowers on the trunk of Epicharis parasitica , an example of cauliflory
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Some members of the Espeletia genus exhibit a growth habit that is caulirosulate .
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Moehringia growing as a chasmophyte on an overhanging cliff
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Chloroplast s within the cells of the leaves of the moss Bryum capillare
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Not all chloroplast s are simple in shape. Chloroplasts of Spirogyra are helical within the tubular cells of their algal filaments.
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The so-called "fleshy leaves" of cacti, such as on this Opuntia tomentosa , are actually cladode s (branches). The true leaves are the spine s growing on the cladodes, which on this young cladode are still fleshy.
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Colony of cells forming a coenobium , of an alga in the genus Pediastrum
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Asclepias syriaca seeds, showing the coma of hairs in its pappus
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Pfaffia gnaphalioides flowers with basal coma hairs
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Coma atop Muscari armeniacum , bearing sterile flowers
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The conical compound inflorescence of Aeonium arboreum is a compound panicle composed of minor panicles, some of which are compound in their turn.
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California buckeye ( Aesculus californica ) has a compound palmate leaf, the leaflets radiating from a central point.
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The lobes of the gamopetalous corolla of Nicotiana flowers are conduplicate in the bud.
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Casuarina equisetifolia male and female flowers and cone s
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Gamopetalous Watsonia flower split open between two petals to show the connate formation of the corolla tube; compare the adnate attachment of the stamen bases to the matching petals
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Corm s , one entire in its tunic , one partly peeled to show tunic cataphyll s, and one split to show inner structure
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The corona of this Passiflora flower is a ring of purple filament s between the petal s and the stamen s.
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Cotyledon s of seedlings of Koelreuteria . One plant shows the first new leaves above its cotyledons, and the rest show various younger stages of emerging cotyledons.
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Crassula rupestris frequently grows as a cremnophyte on cliff faces in fynbos .
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Mimetes cucullatus , so named for the hooded, cucullate shape of its white flowers
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Murraya paniculata has leaves with cuneate (wedge-shaped) bases.
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Cuspidate leaves of Diplacus bigelovii var. cuspidatus
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Euphorbia milii is commercially grown for the aesthetic appearance of its brightly colored, bract -like structures called cyathophyll s , which sit below the inflorescence .
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Seasonal, healthy decortication of Eucalyptus grandis outer bark
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A decorticating machine collecting fiber from leaves
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Astragalus austriacus is regarded as diadelphous because it has one stamen unattached to the main adelphia (bunch).
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The paired cotyledon s of a castor bean seedling ( Ricinus communis ) are typical of a dicotyledon .
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Discolorous leaves of Brachylaena discolor differ in color between their upper and lower surfaces.
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Disk florets opening in a capitulum of a cultivated Helianthus . They open progressively from the edge to the center of the disk.
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Dissepiment developing in tissue of carpels where they meet to form locules in the capsule of the ovary of Lilium
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Boophone disticha has conspicuously distichous leaves.
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Domatia at the bases of the thorns of Vachellia drepanolobium , the whistling thorn, with visible access holes
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Leaves of Epipremnum aureum (golden pothos) have a cuspidate drip tip .
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Plants of the genus Corydalis bear seeds with attached elaiosomes , which have various functions, commonly attracting ants. On some Corydalis species, elaiosomes that attract ants also repel mice. [ 32 ]
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Ficus lyrata is an example of a doubly- emarginate leaf with lateral and apical emargination ; it also might be seen as a basally emarginate .
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Petals of Heracleum sphondylium are variously emarginate at their tips. Flowers in the middle of the inflorescence have slightly emarginate petals, whereas flowers at the periphery are so deeply emarginate as to be almost cleft in two.
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The pale embryo emerging from the upper surface of the sprouting date seed is tiny in comparison to the endosperm , its main food supply, which comprises almost all of the rest of the seed.
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Water lilies and reeds represent two ecological categories of emergent aquatic vegetation .
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Iris pseudacorus has clearly ensiform leaves: narrow, straight-edged, sword-shaped.
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The enlarged calyx and smaller epicalyx of Hibiscus sabdariffa
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Tillandsia recurvata growing as a harmless, non-parasitic epiphloedal epiphyte on a tree trunk that is also infested with an epiphloedal foliose lichen
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Seeds or fruits are dispersed by epizoochory when they stick to the fur of animals.
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The bases of equitant leaves enclose later leaves on the stem.
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Sections of exalbuminous seeds
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Aloe marlothii flowers with stamens and stigmata of mature flowers exserted from the mouths of the floral tubes
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Astragalus falcatus has conspicuously falcate pods; not many falcate anatomical structures are so markedly curved.
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Rhigozum obovatum bears its leaves in well-defined fascicle s .
Trunks and branches of some species of poplars contribute to the trees' fastigiate habit .
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Favolaschia calocera , the orange pore fungus, has conspicuously faveolate fruiting bodies.
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Emerging leaves of Oldenburgia grandis are heavily felted .
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In the wild, the leaves of Fenestraria commonly are covered in soil, except for the transparent fenestration ; this permits photosynthesis while reducing damage from exposure to intrense sunlight and herbivores.
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Digitalis ferruginea owes its specific name to its ferruginous (rust-colored) flowers.
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The pseudanthium of Zinnia elegans is typical of many Asteraceae in that it includes two types of floret s , ray florets and disk florets .
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Medicago sativa ( alfalfa or lucerne) is an agriculturally important forb , grown in large volumes for forage, soil improvement, and other purposes.
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Galbulus (berry-like, fleshy) cones on the coniferous tree Podocarpus elatus
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Gametophores (red male antheridia and brown female archegonia) borne on a gametophyte of a Chara species of green algae
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Longitudinal section of immature male pine cone, showing male gametophytes (pollen grains) developing between the cone scales
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The leaves, buds, and young stalks of Eucalyptus macrocarpa are glaucous , covered with a thick waxy pruinosity .
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Glochids at the base of an Opuntia cactus spine
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Glumes of a grass species with a fairly large inflorescence
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Scanning electron micrograph of a stoma on the leaf of Haemanthus . The two lip-shaped cells on either side of the pore are the guard cells .
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Drops of guttation fluid on the dentate points fringing the immature leaf of a grapevine
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Epidermal hairs on plant leaves
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Multicellular hairs on the edge of a sepal of Veronica sublobata
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Haplostemonous arrangement of stamen s and petal s
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The swollen haustorium of Viscum capense renders the end of the branch stunted compared to the lower part of the branch.
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The fruit of Poncirus is a typical hesperidium .
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Heteroblastic growth is common in Eucalyptus species with leaves that are isobilateral in the mature tree; they generally start life with dorsiventral leaves. Some of these saplings are in the transient stage in which they have both forms of leaves, dorsiventral on lower branches, and isobilateral above.
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The hilum contrasts conspicuously with the rest of the testa in the seeds of many species. In the case of Erythrina species, the colors may be a warning that the seeds are poisonous.
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A hypocarpium forms below the fruits of Sassafras albidum .
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Flowers, fruit and propagule of a Rhizophora "mangle" or mangrove . The apparent root of the propagule is in fact meristematic tissue developing from the hypocotyl . The new plant develops largely from this tissue, especially if it has successfully penetrated into mud in which the new plant can establish itself.
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Imbricate protective cataphylls on dormant buds of Quercus robur
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Petals of Mespilus germanica are imbricate before the flower opens.
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Doubly imparipinnate compound leaf of Melia azedarach
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Indehiscent pods of Libidibia ferrea ; unlike most Fabaceae species, the plant depends on the pods being crushed by large ungulates to disperse the seeds.
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The leaves of Syagrus palms are ' induplicately folded, in contrast to many other palm genera with reduplicate leaves.
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Stamens of Calotropis gigantea are inserted at the base of the corolla.
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The intramarginal veins near the margins of this leaf are outlined in white.
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Two of these three green Asteraceae involucres encase unopened flower heads, and the third supports the open colorful head of emerging flowers. The imbricate phyllaries around the heads of this Malacothrix coulteri suggest the keeled scales of a snake, giving the plant its common name: "snake's head".
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This Begonia leaf shows unusual iridescence for a plant.
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Vachellia karroo bipinnate leaf

A. Rachilla
B. Pinnule
C. Jugary glands
D. Juga (plural of jugum )
E. Base of petiole
F. Petiolary gland
G. Rachis
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Jugate leaf of Bauhinia glabra
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One form of the kettle trap s of a pitcher plant
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Typical knee at a node in a grass stem
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Unusually dense stand of cypress knees around the parent tree
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Laciniate , deeply incised, leaves of Pelargonium crispum
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Most Euphorbias are laticiferous and instantly exude latex when even mildly punctured.
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A leaf scar on Juglans regia , showing the layer of corky protective tissue that remained after the leaf separated along the abscission zone . It also shows the leaf traces of the vascular bundles that broke off when the abscission zone failed. The axillary bud associated with the leaf shows just above the scar.
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The dark horizontal lines on silver birch bark are lenticel s .
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Lignotuber s of Lambertia formosa growing sprouts after a bush fire
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Ligule between the leaf sheath and leaf of a grass
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Loculicidal dehiscence of a fruit capsule . The locule walls split at the back, and the valves separate, bearing the septa on their centers.
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The loment (or lomentum ) of Hedysarum occidentale splits into single-seeded segments along the visible lines of weakness when ripe.
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Maculate leaves
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Eucalyptus socialis , showing its mallee habit , a single tree with several trunks growing from an underground lignotuber
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Mast from beeches on the forest floor
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Apical meristem in root tip:
1: Meristem
2: Columella
showing statocytes with statoliths
3: Lateral part of the tip
4: Dead cells
5: Elongation zone
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Mesophyll as seen in the cross section of a dicotyledonous leaf
A-Lower epidermis
B-Lower palisade mesophyll
C-Upper epidermis
D-Upper palisade mesophyll
E- Spongy mesophyll
F-Leaf vein
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Longitudinal section of Pinus ovule
A= Gametophyte
B=Egg cell
C= Micropyle
D=Integument
E= Megasporangium
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A germinating date palm , Phoenix dactylifera , a monocotyledon , showing its single cotyledon
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Mucro at the tip of the rachis of a compound leaf of Vachellia karroo
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Fungal mycelium grown in culture dish
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Mycelium of mycorrhiza growing on the roots of Picea
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Hoya carnosa secretes so much nectar that it falls in drops if no pollinators remove it.
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The small green petals of Helleborus argutifolius act as floral nectaries . The sepal s function as petal s.
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Some Senna species have extrafloral nectaries that attract ants to defend them from pests.
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Plant stem node s and internodes
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Obovate leaflets of a ternate leaf of Kummerowia
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Obtuse leaves of Dovyalis zeyheri
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Open flower of Eucalyptus macrocarpa , next to a shed operculum
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Opposite arrangement ( phyllotaxis ) of leaves
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Ovate leaflets on a ternate leaf of a Vigna species
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The thick trunk of Brachychiton rupestris accumulates moisture as a means of survival of droughts, and presents a marked example of a pachycaul habit .
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This Curio articulatus is pachycladous in that it has a disproportionately thick stem.
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A maple ( Acer platanoides ) leaf has palmate venation, as its veins radiate out from a central point, like fingers from the palm of a hand.
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Asclepias physocarpa shedding seeds, each with its silky pappus
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Doubly paripinnate leaves of Delonix regia
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Stephania japonica is a vine with peltate leaves.
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Perfoliate leaves of Smyrnium perfoliatum with stems passing through them
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The perigonium of a moss (red in this case), also called a splash-cup , surrounds the antheridia and aids in dispersal of sperm.
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Liquidambar styraciflua bud emerging from its protective brown imbricate cataphyll scales, also known as perule s
Pelargonium lobatum inflorescence , with showy petals projecting from inconspicuous protective sepal s
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Petiolary glands on the petiole of a cherry leaf
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Rock-splitting roots of the petricolous large-leaved rock fig, Ficus abutilifolia
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The phaneranthous habit of the red flowering gum, Corymbia ficifolia , can attract pollinators such as the honey eater, Anthochaera chrysoptera , from a considerable distance.
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Seedlings of Acacia fasciculifera bear leaves that illustrate the ancestral function of their phyllode s as petioles .
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Pileus of the fruiting body of the fungus Pluteus admirabilis
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Glandular pilose hairs on the stem of Aquilegia grata
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Bipinnate leaf anatomy showing a pinna (or pinnule)
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Pistillate flower s of Shepherdia canadensis .
Compare staminate flower .
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Electron micrographs of sections of wood of a conifer ( Picea abies ) show pit s in the tracheid walls.
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Flowers in the inflorescence of Euphorbia platyphyllos open simultaneously, as a pleiochasium .
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Longitudinal section of maize kernel (scale=1.4 mm):
A= pericarp , B= aleurone , C= stalk , D= endosperm , E= coleorhiza , F= radicle , G= hypocotyl , H= plumule , I= scutellum , J= coleoptile
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Pneumatophore s on a species of mangrove
Pollinium of an orchid, stuck to a syrphid fly that visited one flower, and may yet deposit its burden onto another flower of the same species.
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The sharp projections on the trunk of the knobthorn, Senegalia nigrescens , are prickle s rather than thorns, botanically speaking.
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Procumbent growth habit of Sagina procumbens , growing mainly along the soil surface, but without rooting
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Carpobrotus and other prostrate plants growing on sand in Sicily, striking root and binding the soil as they grow
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Floral stages of the protandrous species: Geranium incanum . The flower at first has intensely colored petals, and both androecium and gynoecium . After a day or so in bloom, it sheds the stamens and the color of the petals becomes somewhat paler.
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Punctate glands on Artemisia nova are visible because they are not covered with epidermal hairs .
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Punctiform glands on the undersurface of a Plectranthus leaf
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Pyriform syconium ("fruit") of domestic fig
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Siliques of Conringia orientalis tend toward a quadrate cross section.
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Bulbinella latifolia raceme s . The flowers are already open at the bottom; at the top, the axis is still growing and budding.
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Rachis of Vachellia karroo bipinnate leaf, with components labelled as follows:
A. Rachilla (the diminutive of rachis )
B. Pinnule
C. Jugary glands
D. Juga (plural of jugum )
E. Base of petiole
F. Petiolary gland
G. Rachis
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Radicle s emerging from germinating seeds
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Reniform kidney bean seeds
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The central leaflets of the ternate leaves of Searsia glauca are oblate and commonly retuse .
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Typical rhizome . This one is a specimen of Iris pseudacorus .
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Grafting kiwifruit vine scion onto rootstock below
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Unidentified Gasteria bearing leaves with a rugose surface, banded with callosities
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The runcinate lobes of a Taraxacum officinale leaf point downward, i.e. toward the stem.
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Sagittate leaves of an Alocasia plant
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Trametes versicolor , the turkey tail fungus, is a saprotroph that consumes dead wood in forests. Its common name comes from the conspicuously patterned brackets, but the main body of the saprotroph consists of the largely invisible mycelium that penetrates the dead wood and digests it.
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Strawberry plants reproduce mainly by sarment s , stolons such as these, often called runners ; at their nodes the sarments put up tufts of leaves and strike root if there is any good soil beneath.
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Micrograph of the scabrid undersurface of the leaf of Stipa pulcherrima .
Amaryllis belladonna in flower, an example of a leafless scape emerging directly from the underground bulb before the seasonal leaves
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Involucral bracts of Syncarpha species are as scarious as tissue paper, but look like live petals for years, so they are known as "Everlastings" and valued for dried arrangements.
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Isolated sclereid or stone cell in plant tissue
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Sclereids in gritty particles of pear tissue
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Sepal s on Geranium thunbergii , five separated behind the petal s of an open flower, and a connected set enclosing an unopened bud
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Sericeous leaves of Podalyria sericea , the silver sweet pea bush
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The fruits of Lepidium bonariense are silicles , green and circular, with a notch at the apex.
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Silky foliage of the silvertree, Leucadendron argenteum
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Sori under the leaf of the fern Rumohra adiantiformis . Some are still covered by their indusia .
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Spadix of Amorphophallus maximus within its spathe . The female flowers are around the bottom of the spadix, the male flowers above, and the sterile top part is the major source of pollinator attractants.
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Drosera spatulata leaves are markedly spathulate .
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The flowering spike of this Salvia nemorosa differs from a raceme in that the flowers are practically sessile .
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Spine s emerging from the areoles of an Echinopsis species
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Spinescent leaves of Salsola australis : stiff, narrowed, and with lobes ending in spiny points
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Bird nest fungi, Nidulariaceae , bear examples of splash-cup s with spores that are spread by raindrops.
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Staminate flower s of Shepherdia canadensis
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Manilkara hexandra flowers have both stamen s with anther s and staminode s that have no anthers.
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Subulate leaves are narrow with an elongated, tapering tip, as seen on this species of Aloe .
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The large, succulent , acaulescent , linear , cuspidate mottled leaves of a Gasteria species and the small, succulent, cordate leaves of a Crassula species contrast with the linear , herbaceous leaves of a Hypoxis species.
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Sucker s around the trunk of Dypsis lutescens
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Sulcate (specifically polysulcate) grooves along the stem of Scorzonera cana
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Superior ovary ovary in an Aloe species. One flower is sectioned to display the pistil and hypanthium .
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The suture along the concave curve of the pod of a Crotalaria incana , along which the seeds are attached, is where the single carpel has folded shut.
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An undamaged syconium of a Ficus species, plus two more cut open longitudinally to display the fruit within
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Umbo in the middle of the cap of Cantharellula umbonata
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Thorny prickle s of Senegalia mellifera subspecies detinens are unciform .
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Pitchers of the species Nepenthes ventricosa tend to be markedly urceolate .