[1] These chromosomes, carrying genetic information, align in the equator of the cell between the spindle poles at the metaphase plate, before being separated into each of the two daughter nuclei.
[citation needed] In metaphase, microtubules from both duplicated centrosomes on opposite poles of the cell have completed attachment to kinetochores on condensed chromosomes.
[3] This even alignment is due to the counterbalance of the pulling powers generated by the opposing kinetochore microtubules,[4] analogous to a tug-of-war between two people of equal strength, ending with the destruction of B cyclin.
Any unattached or improperly attached kinetochores generate signals that prevent the activation of the anaphase promoting complex (cyclosome or APC/C), a ubiquitin ligase which targets securin and cyclin B for degradation via the proteosome.
For classical cytogenetic analyses, cells are grown in short term culture and arrested in metaphase using mitotic inhibitor.